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Galvatronus Prime was formed when Galvatron was created from the remains of Megatron.

Megatron's mind was split, and Galvatron became wholly evil, with Galvatronus becoming wholly good. Galvatronus stayed inside Unicron, until he came across Hot Rod battling Galvatron. Galvatronus aided Hot Rod, and opened the Matrix with him, becoming Galvatronus Prime.

Afterwards, Galvatronus, Purify and Hurricanus set off into the universe to attempt to find the Heart of Primus, which is said to be able to restore peace to Cybertron.

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